In the pilot of Smash we are introduced to Tom Levitt and Julia Houston, a song-writing duo who had planned on taking a break from the business of Broadway but, find themselves drawn back in when they get the desire to produce a musical based around the life of Marilyn Monroe. When a demo of one of the tracks goes viral the project attracts the eye and ear of tenacious producer Eileen Rand and, although causing a conflict that of director Derek Wills. But the biggest and most important task is to find their Marilyn after, auditioning a range of girls they whittle it down to two but, who do they go for. Ivy Lynn a veteran of the stage who wants to break away from the chorus line to get her break as a lead or Karen Cartwright a fresh talent who is trying to follow her dream of becoming a star.
Ivy and Karen jump through hoops for the brilliant but temperamental director Derek Wills as they pursue the role of a lifetime. Meanwhile, Julia and her husband Frank, are frustrated by the red tape of an international adoption process, and Eileen fights to finance her new musical in the midst of an acrimonious divorce.
Karen finds herself seduced by the glamorous life of new BFF Rebecca Duvall -- but can Rebecca be trusted? What will Karen's ascent mean to her relationship with Dev? And will Rebecca drive Derek insane? Eileen introduces new beau Nick to ex-husband Jerry, while Julia panics when her son Leo goes missing.
"Bombshell" faces its first audience just as a major crisis hits Rebecca Duvall. The team of Tom and Julia scramble to fix the show when the first preview falls short, and Frank must adjust to the unwelcome return of Michael Swift to his life. Dev seeks reconciliation with Karen, but explosive information about Ivy lurks just beneath the surface.
Tom and Julia work to save the show, while Derek makes a decision that will forever change Karen's and Ivy's lives; Ellis tries to save the show.
Back from a successful run in Boston, Eileen, Julia and Tom are blindsided by both personal and professional obstacles as they seek to take "Bombshell" to Broadway. Karen consults her new mentor, Broadway star Veronica "Ronnie" Moore, as she finds herself on the brink of fame, but her success may come at cost to Ivy. A new talent is discovered in an unexpected place. Meanwhile, Derek copes with a damaging accusation.
Partnerships are tested as Eileen asks Tom and Julia to contend with an unwelcome new collaborator, while Jimmy and Kyle prepare to present their work for the first time. Derek attempts to get Ronnie back on his side. Ivy finds it hard to move on, but help from an old friend might take her one step closer to her dream.
Derek calls in help for a special event, but Jimmy's pride might get in the way of a huge opportunity Karen orchestrates for him and Kyle. Peter challenges Julia's pride as they continue to work together. Meanwhile, Ronnie struggles to assert her independence from a powerful mother, as Eileen finds herself at a moral crossroads with huge ramifications for "Bombshell."
After Leigh Conroy joins the cast of "Bombshell," Tom finds himself struggling to diffuse the years of tension between her and Ivy. Jimmy and Karen’s connection is threatened by both Ana’s newfound success and an unexpected visitor. Derek learns more about Jimmy’s past just as the public sees their first piece of "Hit List."
Tom plans a surprise for Ivy in an attempt to find a balance between their work life and their friendship. Relations between Karen, Jimmy, and Derek explode just as "Hit List's" rehearsal process nears its close. While Richard asks Eileen to spend less time at work and more time with him, Julia finds herself also pulled away from "Bombshell" by an unlikely source.
It's "Bombshell's" opening night. As Ivy's nerves get the better of her, support comes from an unlikely source. Meanwhile, Tom and Julia look to their next project. "Bombshell's" success may be in jeopardy as Eileen's issues with Richard come to a head. Jimmy finally opens up to Karen about his past, but is it too little too late?
An off-the-rails Jimmy threatens "Hit List's" future, as Karen, Derek, Ana and Kyle try to help him before something bad happens. Tom and Julia's partnership is strained to the breaking point just as Ivy, Eileen, and the entire "Bombshell" team are running themselves ragged to outshine the competition.
With his sights set on the Tony, Tom pulls out all the stops at a Houston & Levitt tribute night, but can Julia turn away from "Hit List" long enough to help? Ivy is afraid of damaging her reputation when Tom asks her to perform a risqué number. The cast of "Hit List" is rocked by a betrayal close to home.