Originally entitled Lowbrow, this pilot episode was shown during Cartoon Network's 2002 Summerfest, wherein eight original Cartoon Cartoon pilots were shown. It was chosen as the most popular among viewers, and Lowbrow was greenlit and retitled Megas XLR. The plot is a shorter version of the first full episode "Test Drive"
In the future, the Earth Coalition is fighting a losing war against the Glorft Imperium, a highly advanced extraterrestrial race with an insatiable hunger for conquest. Skilled 'mech pilot Kiva wants to send a robot named "Megas" back to a crucial part of the conflict, but accidentally sends it too far, and Megas winds up in the hands of couch potatoes Coop and Jamie. Worse, Kiva is stuck in the past with these boneheads, not to mention a Legion of really angry Glorft intent on capturing the Megas, even if they have to resort to using the giant UMD Mech to do it.
A floating alien head named Magnaminous sees Coop's fighting skills and invites him to the Galactic Combat Federation to fight various other powerful robots. Coop agrees, but maybe it would have been better if he had stayed home because, after a couple of fights, Magnaminous reveals he wants Coop to lose so Magnaminous can make a fortune. Coop refuses to accept those terms, so Maganaminous resorts to some dirty tricks to this end.
It's a hot day in the city, and Coop just wants a big old thirst quenching Megaslush. However, he accidentally causes a highly destructive robot named the REGIS Mk. V to fall to Earth. Every time Coop gets close to getting his slush, the REGIS shows up and causes the store to be destroyed. Coop is tired of REGIS' continuous desire of destruction and shop-ravaging, but how is he supposed to stop a robot that gets bigger and stronger every time you smash it?
Coop begins a bad day by accidentally creating a giant cheese monster and has to stop it, but the method he uses gets cheese in Megas's CPU core. Unaware of this, Coop enters Megas in an auto show, and is putting up a very good showing until Megas breaks down and nearly destroys the building. Fortunately, he's able to stop it, but now the Glorft see an opening to attack and invade the city, hoping to capture Megas. But first they need to find the CPU core.
Coop, Jamie and Kiva discover an ancient Ringworld while training. They discover it is actually a giant interstellar library with information on almost every species in the universe. Kiva is excited, because this could contain the key to defeating the Glorft and traveling back to the future. However, this Ringworld is infested with robotic, worm-like, energy loving bugs. Coop, in the end, ends up destroying the entire Ringworld when he destroys the worms' Queen.
In an attempt to fix the television, Coop accidentally causes a blast of concentrated television waves to go out into space and attracts a giant, radio-wave eating monster to Earth. Due to the fact it is eating all the TV-satellites, it is impossible for anyone to watch any TV. Coop needs to stop this thing at all costs, but there's one major snag - how do you stop a planet-sized monster that eats television waves, not to mention that it has a bunch of killer boogers inside its body?
Coop buys a cartridge from a garage sale, thinking it's a video game. In reality, it's an interdimensional prison for the galaxy's most dangerous criminals-the most dangerous of which is Grrkek The Planet Killer. So, naturally, Coop unleashes Grrkek first, but is able to beat him with a cool looking laser that traps Grrkek. However, he proceeds to unleash all the other prisoners (including one that looks like a robotic Gamera), which show their gratitude by trying to destroy the city. Naturally, Coop must clean up another one of his own series of messes.
In order to impress a girl, Jamie borrows Coop's car to take her for a ride. Unfortunately, Warmaster Gorrath arrives piloting a deadly replica of Megas (dubbed Mecha-Megas) who's destroying Jersey searching for Megas; naturally, everyone thinks that the fake is just Coop being destructive. So Coop must use his secondary command bridge to control Megas. However, this method is much more difficult, so Coop does rather badly. Even worse, Jamie accidentally activates the car's connection to Megas, so now he's in control of Megas, but he has no idea - he's just trying to have a date.
A group of crime-fighting teenagers, called the S-Force, think Coop's the ultimate villain, so they come to Earth to try to "stop" him. Coop tries to fight them off and explain he's actually a hero, when darn the luck, Ender, a dangerous intergalactic criminal and The S-Force's Arch Nemesis, shows up to help him. Coop needs to defeat Ender, but the S-Force (and Ender) still don't get it.
Coop accidentally damages a vital part of Megas, the photonic stabilizer. Without it, Megas will go up in a blaze of fiery glory and explode, which will not only destroy Megas, but also, everything within a 100-mile radius. Fortunately, Coop found a junk planet that probably has just about everything. Unfortunately, Varshin the Dealer wants Megas' time drive to complete his own robot and sends various "zombie" robots to retrieve it. Even more unfortunate, Goat has come along for the ride, and is thoroughly annoying Kiva, not to mention he's fantasizing about a personal robot of his own without any idea how to get or build one properly.
An intergalactic bounty hunter arrives, and soon after Megas is impounded due to parking regulations. Coop needs to get it back in order to save Kiva, who is the target of the bounty hunter. Unfortunately, Coop's license has expired, so now he has to pass for a new one at the DMV. As Kiva battles with the evil bounty hunter, Coop has to deal with an even scarier villain - the red tape of the Department of Motor Vehicles.
Coop accidentally destroys a sacred treasure of the Azion Worlds, a robotic, alien empire - the Flame of Azeroth. The Emperor declares whoever captures Megas and bring it to judgement will be named as the new Emperor, so now Coop has to fight off a bunch of Azionites in the deep, dark reaches of the final frontier, all of which are, if necessary, willing to destroy each other to become the Emperor. (Note: due to a tape mixup, this episode was not shown until nearly a year later).
Coop accidentally teleports himself directly into the Karrajor, the Glorft Mothership, and now Kiva and Jamie have to figure out how to pilot Megas to save him. They do okay at first, but eventually crash into the Glorft mothership and get themselves - and Megas - captured. Fortunately, Gorrath foolishly lets Coop get the car, so Coop tears through the ship, intent on rescuing his friends and Megas. Now Coop must rescue everyone while finishing off the Glorft invasion once and for all, but how is he supposed to do that when the sheer amount of Glorft mech's overpowers even Megas and tears off several vital parts? Even worse, the Glorft have a new mech, the Karrajor in Mech mode, which is even larger than the UMD, which raises the question: how will Coop defeat this new threat with a critically damaged Megas?
In an effort to impress some Sailor Senshi-style space girls, known as the Ultra-Cadets, Jamie poses as Coop and is taken to their planet, where he learns the Ultra Cadets' city is in trouble from a giant fire monster. Armed only with an age-old, feminine robot, Jamie "fights" the monster and defeats it the way Coop always does: sheer luck. However, Coop arrives with Megas to save Jamie and gets into a fight with the mechanized Ultra Cadets.
It's 5:01 and a video is due back at the rental store by 5:30, and Coop has to get it there on time to keep his membership from being revoked. However, Magnanimous appears and challenges Coop to protect his title against several robots, including one resembling Mr. T. Coop manages to beat the robots, but inadvertently challenges the entire galaxy to fight him - and the galaxy takes him up on the offer.
Coop inadvertently frees the Glorft from no-space, allowing Gorrath to start his revenge. The only thing Coop is worried about, however, is babysitting cousin Skippy, who is easily bored and hates everything about Coop's house. In an effort to get him to shut up, Coop takes Skippy to the moon in Megas. (Incredibly, he is still bored.) There he discovers the Glorft is plotting to smash the moon into the Earth. Coop has to fight off the Glorft while making sure Skippy doesn't end up Glorft fodder.
Coop, Jamie and Kiva head off to Las Vegas, and inadvertently discover Area 50. Coop accidentally activates a destructive energy-leech robot, who, due to severe and fatal programming errors, sees everything as "the enemy." Coop further screws up by directing the robot to Las Vegas. Now Coop must find a way to shut down the robot permanently before Las Vegas loses all its power. Coop in the end destroying the cities power source causing the robot to deactivate. And he then buried the robot in the Grand Canyon which he had to fill with dirt.
Once again, Coop mistakenly releases a giant, plant-type alien menace that wants to destroy Earth and acquire its heat. Fortunately, Megas is too big for the plant to handle, so it has to find a bigger form. Unfortunately, this is Thanksgiving, and the spore fuses with a bunch of parade balloons to this end. Coop now must prevent the plant from wrecking the holiday.
The S-Force has been captured by an evil villain named Zarek, and he is planning to execute them. Fortunately, Jax escapes and manages to enlist Coop's help. Coop manages to save the S-Force from the "infinity zone," which Coop, Jamie, Kiva, the S-Force, and Megas fall into, but survive the Infinity Zone after Coop helps everyone escape, but now Zarek is sending down an entire army to terminate them. It's Megas and the Ultra-Dimensional Power Zorp vs. a bunch of insect robots! The "gift" that Zarek offers Coop, Jamie and Kiva is an end to their lives, whereas the "gifts" Coop tries to give Zarek, but almost destroys the S-Force with, is actually thousands of missiles being fired at Zarek, due to the new Super Destructor Mode.
Kiva gets some unwanted attention from Captain Warlock, a space pirate who goes for red heads, and now must make a choice between a possible time drive or two complete idiots. As usual, one wrong choice means certain death. Now Kiva must find a way to free Coop and Jamie as well as fend of the advances of one love sick space pirate and his mechanized crew.
The Glorft, disguised as a motorcycle gang thanks to their latest Holo-Camouflages, comes to a rock concert. They're not there for entertainment, however - they want to eliminate Kiva's ancestor, which in turn would cause Kiva to cease to exist, which in turn would mean Megas would stay with the Glorft and never reach Coop, which would in turn doom Earth into extinction. Kiva and Jamie must protect the ancestor until Coop can get to Megas. To make matters worse, it seems that Jamie and has fallen for Kiva's ancestor and she has fallen for him.
Coop crashlands on an icy planet and ends up destroying its only mechanized guardian. Now he must protect the Yetis there from selfish, ice-hungry robots named the Cerilians, who use it to maintain their cooling systems. Trouble is, Megas is iced over, and does some of the Cerilian Army's work for him. This is going to be one icy battle for the team.
Coop, Jamie and Kiva teleport themselves to a planet with brainwashed robots who act like sissies and workers. The alien who did it tries to do the same to Megas, but his brainwasher device only works on sentient robots. (Obviously, Coop's doesn't count.) Coop tries to put the alien out of business for good without unnecessarily destroying the prisoners. Coop in the end destroys the mind control device, freeing the robots. He then leaves the planet, unfortunatly as the group leaves it's shown that the planet was actually a prison to the worst robots in the universe. Who start destroying the planet after they leave.
Coop's builds the world's most powerful universal remote - and Skalgar (who Coop and Jamie called "School Girl"), an ignorant, alien criminal with an inferiority complex, wants it, believing its (non-existent) destructive power can boost his reputation. Coop underestimates his short opponent, and Skalgar teleports the remote to his own mech. Coop then typically destroys half of Jersey City attempting to recover the remote.
In the last days of the Earth War, Jamie and Kiva were busy shutting down the Glorft Core Destroyer, while Coop and the Glorft, led by Gorrath as usual, were fighting out. However, Coop and Gorrath get transported to a mirror dimension - because of a gamepad combo that activated the Transdimension Device - where Coop meets his alternate self, a futuristic, evil, and muscular warlord version of himself at that, having defeated the Glorft, abandoned Megas, formed an Empire, and keeps on destroying - no matter the location, time, dimension and enemy. He also sees that Kiva has turned cyborg as well and is evil Coop's sidekick. Alternate Jamie, on the other hand, has lost his cowardice forever and became strong enough to try and defeat evil Coop and his empire with his Resistance (and on a side note, Jamie and Kiva were dating in the future). Coop and Gorrath must work together this time to stop mirror Coop and prevent him from ever destroying Earth again.
In an attempt to stop evil Coop from destroying their dimension, Coop and Gorrath, assisted by alternate Jamie, launch a desperate assault on the evil Coop. Tragically, Megas is (once again) critically damaged and Coop (in a suicide charge) gambles everything in attempting to stop the Evil Coop but only succeeds in getting his core critically damaged and causing Megas to finally be destroyed, but the protagonists suddenly realize that evil Coop abandoned his Megas in favor of his more warlike mech. Then evil Coop and his massive Grunt army invades Coop's dimension, and both of them swiftly create a giant battlefield, starting the big but short Dimensional War. On the Evil side, is evil Coop in his Black and Red Warlord Mech with hundreds of mindless Grunt 'Mechs and evil Kiva in her separate EC Mech, while the Good side, contains Coop in evil Coop's MEGAS, Gorrath in his personal Mech, including his army of Glorft Grunt 'Mechs. After a long and devastating battle that reduced the evil Coop's army into mere junk, mirror Coop and mirror Kiva retreat into the Dimensional Gate to return to his dimension. As soon as the mirror leaders entered the Dimensional Gate, Coop orders alternate Jamie (Who remained in evil Coop's stronghold) to destroy the Gate, trapping Evil Coop and Evil Kiva stuck between worlds forever, never to cause annihilation of any dimension again. (In this case, it was a world filled with Lerps - elf-like creatures similar to The Smurfs, but they can breathe flowers and go angry when attacked.) Meanwhile, Coop decides to test what new gadgets the evil Coop installed into Megas and destroying the city once more in the process.