Tom and Twigs find Rickety's old racing car, the Turbo Rantula, and challenge the Mushas to the most famous race in Treetopolis, the Conkerball Run. They compete to win back the Conkerball Cup, a prize Rickety lost to the Mushas a long time ago. Tom, Twigs and Zigzoo form Team Treeling and learn to work together to overcome epic obstacles and cheating Mushas. It's a race to the finish with team work the winner.