Young cub Pip's adventurous nature soon gets him into trouble as he wanders away from the pride. A lost, lonely young lion is easy prey for prowling leopards and hyenas. Meanwhile Pip's cousins Ziggy and Prince have troubles of their own. They've survived an attack from a rival male lion, but now their mum Rosa is trying to re-join her pride. Ziggy and Prince's new step-dads are far from happy.
The prides have overlapping territories; which means the action and events in one lion family unavoidably ends up affecting the other. The discovery that ‘Ava', one of the dwindling Hollywood Pride, has a litter of new-born cubs hidden away is great news. However, the trio of ‘Punk' Males that lead the neighbouring pride are expanding Hollywood territory, putting the cubs in immense danger.
It's now heading towards the rains, and for young cubs Prince and Ziggy it seems that they may finally have a chance of surviving this perilous and eventful season; against all odds they've been accepted into the pride by their step-fathers. But a jealous aunt takes exception to their presence in the most brutal way, and the brothers face tragedy.