With the reappearance of a freighter ship from 1963, the A.E.G.I.S. Far-East branch is greeted by another special Invader, calling himself Kikai Shogun (General Machine). With the ability to merge with machines, the general chooses the Hikari bullet-train as his next test subject.
Riding this train is Shun and his family, who are returning from a brief vacation. With the train on a full speed coarse to Tokyo Station, the rest of the Gate Keeper team must intercept it before it crashes. Either the train must be stopped before it reaches the station, or be destroyed!
Un des leaders des envahisseurs, le Méca-Général, a l'intention de faire dérailler un Shinkansen, le train ultra-rapide. Les Gate Keepers entrent en action, mais la situation se complique quand ils apprennent qu'à son bord Shun est détenu en otage par le général.