Edward Kynaston (Billy Crudup) is one of the leading actors of his day, particularly famous for his portrayal of female characters, predominantly Desdemona in ''Othello''. His Dresser (theatre), Maria (Claire Danes), aspires to perform in the legitimate theatre but is forbidden because of a law enacted by the Puritans prior to the restoration of the House of Stuart. Instead, she appears in productions at a local tavern under the pseudonym Margaret Hughes. Her activity is aided by the novelty of a woman acting in public, which attracts the attention of Sir Charles Sedley (Richard Griffiths), who offers his patronage. Eventually she is presented to Charles II of England (Rupert Everett).
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